Gagea shmakoviana Levichev

Accepted Name:

Gagea shmakoviana Levichev, 2001. Turczaninowia, 4 (1-2): 28, tab. 8; id. 1999. Komarovia, 1: 54, nom. nud.

Description

Plantae virides, mollifoliae, solitariae, 15 21 cm alt. Bulbus 7 9 mm in diam., oblique guttiformis, tunicis fuscis tenuiter coriaceis tectus. Plantae juveniles bulbillo solitario, plantae immaturae bulbillis paucis, quorum basalis et eo oppositus sororiis reliquiis majoribus, bulbilli vegetativi duo in statu generativo ineunte ab utroque latere ad basin pedunculi formati, bulbilli globosi, 1.5-3 mm in diam., minute verruculosi, atro-fusci. Pedunculus 10 15 cm altus, sectione 6 5 4-angularis, in statu generativo ineunte 6-angularis bicanalisque, in statu praesenili 4 5-angularis unicanalisque vel 4-angularis canalis nullis. Folium radicale solitarium, inflorescentia sublongius, superne reflexum, 2.5 4.5 mm lt., anguste lineare, sectione late angulosum. Folia caulina 2 4(5), verticillata, quorum infimum e radicalibus enatum, inflorescentia brevius vel ei aequilongum, ca 3 4(5) mm lt., anguste lanceolatum. Inflorescentia 2 3(5)-flora, umbelliformis, pedicellis 2.5 4 cm lg. Perianthii phylla 13 15 mm lg., 1.5 2 mm lt., anguste lanceolata, apice obtusata, intus aurea, extus viridia, exteriora interioribus longiora et latiora. Ovarium ovoideum, sessile. Antherae flavae, oblongae (3 3.5 mm lg.), dehiscentes subglobosae (1.5 mm lg.). Capsula obovoidea, perianthio subduplo minor. Semina teretia.

Typus

Rossia, Provincia Altai, districtus Charyschensis, in viciniis pagi Sentelek, 3 km adverso flumine Sentelek, 51 12' l. b., 83 42' l. o., declive expositionis australis, 20.V.1996, A. I. Shmakov, G. A. Terechina, N. V. Elesova, E. Antoniuk, O. Torczakov (LE, isotypus SSBG).

Habitat

Habitat in declivibus apertis herbosis, inter frutices, ad margines agrorum, solo subtiliter argilloso in promontoriis et regione inferiore montium Altaici.

Nomenclature Note

Species in honorem flore Altaici investigatoris et peritissimi filicum cl. A. I. Shmakovii denominata est.

Distribution

5. Siberian (N)

Image(s)

Fig. . Line Art Image Igor G. Levichev 2000 G.shmakoviana.

Commentary

While the unique(sole) species of a genus, at which is revealed the evident proof of accretion of the present bottom cauline leaf with a peduncle (flower stalk) from a corm(lecus) up to an inflorescence. The second basal leaf at G. shmakoviana also grows together on all an extent with a peduncle (flower stalk) and is separated under an inflorescence, simulating bottom cauline leaf. Last is characteristic for many species of a genus, but the accretion of a cauline leaf is confirmed for the first time, though to indirect attributes, it was supposed and earlier. In this case, presence at axials these leafs of bulblets (fig. 2, 8), results in preservation on all an extent of a peduncle (flower stalk) of one or two channels (fig. 8d, e, 16d, 17d, e - 19d, e, 20d). At teratologycal deviations(rejections) with a untypical rule(situation) of a bulblet in an average part of a peduncle (flower stalk) (fig. 2) the bulblets the channel are lower is absent (fig. 15), the channel is higher - is present and well additional circle of bundles conducting from an accrete leaf (fig. is traced. 16). The fabrics accrete to a peduncle (flower stalk) leafs (second basal and bottom cauline) on fig. are similarly located. 8d, e, 17d, e - 19d, e. Presence of channels and degree of accretion explain the variability, mentioned above and submitted in drawing, of the form of section of a peduncle (flower stalk).
In immature stages without formation(education) of a peduncle (flower stalk) sometimes can develop more than two bulblets (fig. 6, 10). Per the first year generative stages usually there are two bulblets (fig. 7, 11, 12) two channels (fig. also are formed. 8, 17-19). In a late generative stage - one (fig. 2, 14, 16, 20) or any bulblets (fig. 9, 21).